The Joes soon discover that the mastermind behind the terrorist attack is Cobra Commander (Jonathan Pryce), who has teamed up with a wealthy and powerful tech mogul named Rex Lewis (Jon Voight).
Along the way, they team up with Flint (Marlon Wayans), a young and skilled soldier who joins the Joes. The team also meets Roadblock (Common), a skilled hacker and mechanic, and Storm Shadow (Lee Byung-hun), a former ally who has turned against the Joes.
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The movie begins with the G.I. Joes, led by Duke (Channing Tatum), being ambushed by a group of heavily armed mercenaries. The Joes are framed for a terrorist attack on the US Capitol building, and they must go on the run to clear their names.